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Clade
Subclade Archosauria: Avemetatarsalia to Neornithes 

Taxon Feedback
Taxon Arctometatarsalia 
Nominal Author Holtz 1994 
2° Nominal Author  
Taxon Status active
inactive
Comments  
Potential Synonomy Ornithomimiformes, Ornithomimosauria 

Active Phylogenetic Definition Feedback
Active Definition  
Shorthand  
Definitional Author  
Definition Status original
textual substitution
first-order revision
second-order revision
Definition Type node
stem
Node-Stem Triplet yes
no
Other Triplet Taxa  
Specifiers
Specifier A  
Specifier(s) B  
Specifier(s) C  
Qualifiers
+Taxon  
-Taxon  
Datum  
Taxonomic Content stem (entirely extinct)
crown (extant-bounded)
mixed (extant/extinct-bounded)
Publication Year  

Inactive Taxon Status Feedback
Rejection Criteria junior synonym
historically polyphyletic
incomplete definition
no definition
apomorphy-based
redundant
other 
Critique

This taxon is considered inactive, because there is little support for the taxon under its original apomorphy-based definition. Arctometatarsalia was coined as the flagship taxon to reorganize theropod phylogeny (Holtz 1994). Holtz coined and defined Arctometatarsalia as the first theropod with the arctometatarsalian condition and all of its descendants, an incomplete definition (as there is no specific internal specifier mentioned). He intended the clade to include Ornithomimosauria, Tyrannosauridae, Troodontidae, Elmisauridae and Avimimus but to exclude Dromaeosauridae and birds.

Later Holtz (1996) replaced this form-qualified definition with a simple stem-based definition: the clade composed of Ornithomimus and all theropods sharing a more recent common ancestor with Ornithomimus than with birds. Subsequently Holtz (2000) replaced the vulgar term “birds” with Neornithes.

The new taxonimic content of the taxon, however, no longer resembles the original, which was the signpost for a radically different phylogenetic hypothesis within Coelurosauria (see Ornithomimosauria). Because other taxa are available that do not have this history, Arctometatarrsalia is here regarded as inactive.

 
 

Current Age Range Feedback
Earliest Record  
Latest Record  
Range (My)  
Basis for Range  

Definitional History #1
Definition 1 The first theropod to develop the arctometatarsalian pes and all of its descendants. 
Author Holtz 1994:1108 
Type node  stem 
Specifiers (none) 
Qualifiers  

Definitional History #2
Definition 2 Ornithomimus and all theropods sharing a more recent common ancestor with Ornithomimus than with birds. 
Author Holtz 1996:536 
Type node  stem 
Specifiers Ornithomimus, birds 
Qualifiers  

Definitional History #3
Definition 3 Ornithomimus and all taxa sharing a more recent common ancestor with Ornithomimus than with Neornithes. 
Author Holtz 2000:28 
Type node  stem 
Specifiers Ornithomimus, Neornithes 
Qualifiers